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Information Bank for Leptodactylus Frogs  
  
 
 
 
 

 
 
Publications

Amaro-Ghilardi, R.C., G.O. Skuk, R.O. de Sá, M.T. Rodrigues, & Y. Yonenaga-Yassuda.  2006.  Karyotypes of eight species of Leptodactylus (Anura, Leptodactylidae) with a description of a new karyotype for the genus. Phyllomedusa 5:119-133.

Brandão, R. A., & W. R. Heyer.  2005.  The complex calls of Leptodactylus pustulatus (Amphibia, Anura, Leptodactylidae).  Amphibia-Reptilia, 26(4):566-570.

Camargo, A., R. O. de Sá, & W. R. Heyer. 2006.  Phylogenetic analyses of mtDNA sequences reveal three cryptic lineages in the widespread neotropical frog Leptodactylus fuscus (Schneider, 1799) (Anura, Leptodactylidae).  Biological Journal of the Linnean Society, 87:325-341.

Camargo, A., W. R. Heyer, and R. O. de Sá.  2009.  Phylogeography of the frog Leptodactylus validus (Amphibia: Anura): Patterns and timing of colonization events in the Lesser Antilles.  Molecular Phylogenetics and Evolution, 53:571-579.

Caramaschi, U., R. O. de Sá, & W. R. Heyer.  2005.  Common names for the frog genus Leptodactylus (Amphibia, Anura, Leptodactylidae).  Herpetological Review, 36(2):119-120.

Cardoso, A. J., & W. R. Heyer.  1995.  Advertisement, aggressive, and possible seismic signals of the frog Leptodactylus syphax (Amphibia, Leptodactylidae).  Alytes, 13:67-76.`

Crombie, R. I., & W. R. Heyer.  1983.  Leptodactylus longirostris (Anura: Leptodactylidae):  Advertisement call, tadpole, ecological and distributional notes.  Revista Brasileira de Biologia, 43:291-296.

de Sá, R. O.  2005.  Crisis global de biodiversidad:  Importancia de la diversidad genética y la extinción de anfibios. Agrociencia 9:513-522.

de Sá, R. O., & N. Berois.  1986.  Spermatogenesis and histology of the testes of the caecilian, Chthonerpeton indistinctum.  Journal of Herpetology, 20:510-514.

de Sá, R. O., R. Brandão, & L. D. Guimarães.  2007.  Description of the tadpole of Leptodactylus pustulatus Peters, 1870 (Anura: Leptodactylidae).  Zootaxa, (1523):49-58.

de Sá, R. O., & A. Gehrau.  1983.  Observaciones sobre la biología de Phyllomedusa iheringi Boulenger, 1885, (Anura, Hylidae).  Boletín de la Sociedad Zoológica del Uruguay, Segunda Época, 1:44-49.

de Sá, R. O., W. R. Heyer, & A. Camargo. 2005. Are Leptodactylus didymus and L. mystaceus phylogenetically sibling species (Amphibia, Anura, Leptodactylidae)?  ).  In N. Ananjeva & O. Tsinenko (editors), Herpetologica Petropolitana.  Proceedings of the 12th Ordinary General Meeting of the Societas Europaea Herpetologica, August 12-16, 2003, St. Petersburg, Russia.  Russian Journal of Herpetology, 12(Supplement):90-92.

de Sá, R. O., W. R. Heyer, & A. Camargo.  2006 ("2005"). A phylogenetic analysis of Vanzolinius Heyer, 1974 (Amphibia, Anura, Leptodactylidae):  Taxonomic and life history implications.  Arquivos do Museu Nacional, Rio de Janeiro, 63(4):707-726.

Dixon, J. R., & W. R. Heyer. 1968. Anuran succession in a temporary pond in Colima, Mexico.  Bulletin of the Southern California Academy of Sciences, 67:129-137.

Dubois, A., & W. R. Heyer.  1992.  Leptodactylus labialis, the valid name for the American white-lipped frog (Amphibia: Leptodactylidae).  Copeia, 1992:584-585.

Fernández, M., D. Cole, W. R. Heyer, S. Reichle, & R. O. de Sá.  2009.  Predicting Leptodactylus (Amphibia, Anura, Leptodactylidae) distributions:  Broad-ranging versus patchily distributed species using a presence-only environmental niche modeling technique.  South American Journal of Herpetology, 4(2):103-116.

Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green, & W. C. Wheeler.  2006.  The amphibian tree of life.  Bulletin of the American Museum of Natural History, (297):1-37 + insert.

Furness, A. I., R. W. McDiarmid, W. R. Heyer, & G. R. Zug.  2010.  Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae).  South American Journal of Herpetology, 5(1):13-29.

García Pérez, L., & W. R. Heyer.  1993.  Description of the advertisement call and resolution of the systematic status of Leptodactylus gracilis delattini Müller, 1968 (Amphibia: Leptodactylidae).  Proceedings of the Biological Society of Washington, 106:51-56.

Hayek, L. C., & W. R. Heyer. 2005. Determining sexual dimorphism in frog measurement data: Integration of statistical significance, measurement error, effect size and biological significance. Anais da Academia Brasileira de Ciências, 77:45-76.

Hayek, L. C., W. R. Heyer, & C. Gascon.  2001.  Frog morphometrics:  A cautionary tale.  Alytes, 18:153-177.

Heyer, M. M., W. R. Heyer, & R. O. de Sá.  2006.  Leptodactylus fragilis.  Catalogue of American Amphibians and Reptiles, (830):1-26.

Heyer, M. H., W. R. Heyer, & R. O. de Sá.  2009.  Bibliography of the frogs of the Leptodactylus Clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae).  Volume 1.  References.  Smithsonian Herpetological Information Service, Special Publication 1:1-677.

Heyer, M. H., W. R. Heyer, & R. O. de Sá.  2009.  Bibliography of the frogs of the Leptodactylus Clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae).  Volume 2.  Indices.  Smithsonian Herpetological Information Service, Special Publication 2:1-167.

Heyer, M. M., W. R. Heyer, S. Spear, & R. O. de Sá.  2003.  Leptodactylus mystacinus.  Catalogue of American Amphibians and Reptiles, (767):1-11.

Heyer, W. R. 1967. A herpetofaunal study of an ecological transect through the Cordilla de Tilarán, Costa Rica.  Copeia, 1967:259-271.

Heyer, W. R. 1968. The proper name for the type-species of the genus Leptodactylus.  Copeia, 1968:160-162.

Heyer, W. R. 1969. Studies on the genus Leptodactylus (Amphibia, Leptodactylidae).  III.  A redefinition of the genus Leptodactylus and a description of a new genus of leptodactylid frogs.  Contributions in Science, Los Angeles County Museum of Natural History, (155):1-14.

Heyer, W. R. 1969. Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae).  V.  Taxonomic notes on L. latinasus, rhodonotus, romani, and wuchereri.  Herpetologica, 25:1-8.

Heyer, W. R. 1969. The adaptive ecology of the species groups of the frog genus Leptodactylus (Amphibia, Leptodactylidae). Evolution, 23:421-428.

Heyer, W. R.  1970.  Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae).  VI.  Biosystematics of the Melanonotus group.  Contributions in Science, Los Angeles County Museum of Natural History, (191):1-48.

Heyer, W. R. 1970.  Studies on the genus Leptodactylus (Amphibia, Leptodactylidae).  II.  Diagnosis and distribution of the Leptodactylus of Costa Rica.  Revista de Biología Tropical, 16:171-205.

Heyer, W. R.  1971.  Leptodactylus labialis.  Catalogue of American Amphibians and Reptiles, (104):1-3.

Heyer, W. R.  1972.  The status of Leptodactylus pumilio Boulenger (Amphibia, Leptodactylidae) and the description of a new species of Leptodactylus from Ecuador.  Contributions in Science, Los Angeles County Museum of Natural History, (231):1-8.

Heyer, W. R.  1973.  Systematics of the Marmoratus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae).  Contributions in Science, Los Angeles County Museum of Natural History, (251):1-50.

Heyer, W. R. 1974. Relationships of the marmoratus species group (Amphibia, Leptodactylidae) within the Subfamily Leptodactylinae. Contributions in Science, Los Angeles County Museum of Natural History, (253):1-46.

Heyer, W. R.  1974.  Vanzolinius, a new genus proposed for Leptodactylus discodactylus (Amphibia, Leptodactylidae).  Proceedings of the Biological Society of Washington, 87:81-90.

Heyer, W. R.  1975.  A preliminary analysis of the intergeneric relationships of the frog family Leptodactylidae.  Smithsonian Contributions to Zoology, (199):1-55. [This article is broken up into two pdf files.  pdf file 1.  pdf file 2.]

Heyer, W. R.  1975.  Adenomera lutzi (Amphibia: Leptodactylidae), a new species of frog from Guyana.  Proceedings of the Biological Society of Washington, 88(28):315-318.

Heyer, W. R.  1976.  Notes on the frog fauna of the Amazon Basin.  Acta Amazonica, 6:369-378.

Heyer, W. R.  1977.  A discriminant function analysis of the frogs of the genus Adenomera (Amphibia: Leptodactylidae).  Proceedings of the Biological Society of Washington, 89:581-592.

Heyer, W. R.  1977.  Taxonomic notes on frogs from the Madeira and Purus rivers, Brasil.  Papéis Avulsos de Zoologia, 31:141-162.

Heyer, W. R.  1978.  Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae).  Natural History Museum of Los Angeles County Science Bulletin, (29):1-85.  [This article is broken up into two pdf files.  pdf file 1pdf file 2.]

Heyer, W. R. 1979. Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia: Leptodactylidae). Smithsonian Contributions to Zoology, (301):1-43.

Heyer, W. R.  1983.  Clarification of the names Rana mystacea Spix, 1824, Leptodactylus amazonicus Heyer, 1978 and a description of a new species, Leptodactylus spixi (Amphibia: Leptodactylidae).  Proceedings of the Biological Society of Washington, 96:270-272.

Heyer, W. R.  1984.  The systematic status of Adenomera griseigularis Henle, with comments on systematic problems in the genus Adenomera (Amphibia: Leptodactylidae).  Amphibia-Reptilia, 5:97-100.

Heyer, W. R. 1998. The relationships of Leptodactylus diedrus (Anura, Leptodactylidae). Alytes, 16:1-24.

Heyer, W. R.  1988.  On frog distribution patterns East of the Andes.  Pages 245-273 in P. E. Vanzolini & W. R. Heyer (editors), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987.  Academia Brasileira de Ciências.

Heyer, W. R.  1994.  Variation within the Leptodactylus podicipinus-wagneri complex of frogs (Amphibia: Leptodactylidae).  Smithsonian Contributions to Zoology, (546):1-124.

Heyer, W. R.  1995.  South American rocky habitat Leptodactylus (Amphibia: Anura: Leptodactylidae) with description of two new species.  Proceedings of the Biological Society of Washington, 108:695-716.

Heyer, W. R.  1997.  Geographic variation in the frog genus Vanzolinius (Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 110:338-365.

Heyer, W. R.  2000.  The lectotype of Cystignathus hylodes Reinhardt and Lütken, 1862.  Journal of Herpetology, 34:150-153.

Heyer, W. R.  2002.  Leptodactylus fragilis, the valid name for the Middle American and northern South American white-lipped frog (Amphibia: Leptodactylidae).  Proceedings of the Biological Society of Washington, 115:321-322.

Heyer, W. R.  2005.  Leptodactylus fragilis Brocchi, 1877.  White-lipped frog.  Pages 500-501 in M. J. Lannoo (editor), Amphibian Declines.  The Conservation Status of United States Species.  University of California Press:  Berkeley & Los Angeles.  xxi+1094 pages.

Heyer, W. R.  2005.  Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern  South America, and Amazonia.  Arquivos de Zoologia, 37(3):269-348.

Heyer, W. R., & C. L. Barrio-Amorós.  2009.  The advertisement calls of two sympatric frogs, Leptodactylus lithonaetes (Amphibia: Anura: Leptodactylidae) and Pristimantis vilarsi (Amphibia: Anura: Strabomantidae).  Proceedings of the Biological Society of Washington, 122(3):282-291.

Heyer, W. R., & M. S. Bellin.  1973.  Ecological notes on five sympatric Leptodactylus (Amphibia, Leptodactylidae) from Ecuador.  Herpetologica, 29:66-72.

Heyer, W. R., & K. A. Berven.  1973.  Species diversities of herpetofaunal samples from similar habitats at two tropical sites.  Ecology, 54:642-645.

Heyer, W. R., U. Caramaschi, & R. O. de Sá.  2006.  Rana ocellata Linnaeus, 1758 (currently Leptodactylus ocellatus; Amphibia, Anura:  Proposed conservation of usage of the specific name by the designation of a neotype.  Bulletin of Zoological Nomenclature, 63(3):184-186.

Heyer, W. R., & C. M. Carvalho.  2000.  Calls and calling behavior of the frog Leptodactylus natalensis (Amphibia: Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 113:284-290.

Heyer, W. R., J. Coddington, W. J. Kress, P. Acevedo, D. Cole, T. L. Erwin, B. J. Meggers, M. G. Pogue, R. W. Thorington, R. P. Vari, M. J. Weitzman, & S. H. Weitzman. 2000 ("1999").  Amazonian biotic data and conservation decisions.  Ciência e Cultura, Journal of the Brazilian Association for the Advancement of Science, 51:372-385.

Heyer, W. R., & R. I. Crombie.  2005.  Leptodactylus lauramiriamae, a distinctive new species of frog (Amphibia: Anura: Leptodactylidae) from Rondônia, Brazil.  Proceedings of the Biological Society of Washington, 118(3):590-595.

Heyer, W. R., R. O. de Sá, J. R. McCranie, & L. D. Wilson.  1996.  Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae):  Natural history notes, advertisement call, and relationships.  Herpetological Natural History, 4:169-174.

Heyer, W. R., R. O. de Sá, & S. Muller.  2002.   Leptodactylus silvanimbus.  Catalogue of American Amphibians and Reptiles, (742):1-5.

Heyer, W. R., R. O. de Sá, & S. Muller. 2005. The enigmatic distribution of the Honduran endemic Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae). Pages 81-101 in M. A. Donnelly, B. I. Crother, C. Guyer, M. H. Wake, & M. E. White (editors), Ecology and Evolution in the Tropics: A Herpetological Perspective. University of Chicago Press: Chicago & London. xv + 675 pages.

Heyer, W. R., R. O. de Sá, & A. Rettig.  2005.  Sibling species, advertisement calls, and reproductive isolation in frogs of the Leptodactylus pentadactylus species cluster (Amphibia, Leptodactylidae).  In N. Ananjeva & O. Tsinenko (editors), Herpetologica Petropolitana.  Proceedings of the 12th Ordinary General Meeting of the Societas Europaea Herpetologica, August 12-16, 2003, St. Petersburg, Russia.  Russian Journal of Herpetology, 12(Supplement):35-39.

Heyer, W. R., & M. J. Diment.  1974.  The karyotype of Vanzolinius discodactylus (Amphibia, Leptodactylidae).  Proceedings of the Biological Society of Washington, 87:327-336.

Heyer, W. R., J. M. García-López, & A. J. Cardoso.  1996.  Advertisement call variation in the Leptodactylus mystaceus species complex (Amphibia: Leptodactylidae) with a description of a new sibling species.  Amphibia-Reptilia, 17:7-31.

Heyer, W. R., and A. A. Giaretta.  2009.  Advertisement calls, notes on natural history, and distribution of Leptodactylus chaquensis (Amphibia: Anura: Leptodactylidae) in Brasil.  Proceedings of the Biological Society of Washington, 122(3):292-305.

Heyer, W. R., & M. M. Heyer.  2001.  Leptodactylus lithonaetes.  Catalogue of American Amphibians and Reptiles, (723):1-3.

Heyer, W. R., & M. M. Heyer.  2002.   Leptodactylus elenae.  Catalogue of American Amphibians and Reptiles, (742):1-5.

Heyer, W. R., & M. M. Heyer.  2004.  Leptodactylus furnarius.  Catalogue of American Amphibians and Reptiles, (785):1-5.

Heyer, W. R., & M. M. Heyer. 2006. Leptodactylus knudseni. Catalogue of
American Amphibians and Reptiles, (807):1-12.

Heyer, W. R., & M. M. Heyer. 2006. Leptodactylus natalensis. Catalogue
of American Amphibians and Reptiles, (808):1-5.

Heyer, W. R., M. M. Heyer, & R. O. de Sá. 2008. Leptodactylus cunicularius.  Catalogue of American Amphibians and Reptiles, (845):1-5.

Heyer, W. R., M. M. Heyer, & R. O. de Sá. 2010. Leptodactylus savagei..Catalogue of American Amphibians and Reptiles, (867):1-19.

Heyer, W. R., M. M. Heyer, & R. O. de Sá. 2010. Leptodactylus syphax. Catalogue of American Amphibians and Reptiles, (868):1-9.

Heyer, W. R., & F. A. Juncá.  2003.  Leptodactylus caatingae, a new species of frog from eastern Brazil (Amphibia: Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 116(2):317-329.

Heyer, W. R., R. W. McDiarmid, & D. L. Weigmann.  1975.  Tadpoles, predation and pond habitats in the tropics.  Biotropica, 7:100-111.

Heyer, W. R., & L. R. Maxson.  1982.  Distributions, relationships, and zoogeography of lowland frogs:  The Leptodactylus complex in South America, with special reference to Amazonia.  Pages 375-388 in Ghillean T. Prance (ed.), Biological Diversification in the Tropics.  Columbia University Press.

Heyer, W. R., & L. R. Maxson.  1982.  Neotropical frog biogeography:  Paradigms and problems.  American Zoologist, 22:397-410.

Heyer, W. R., & V. R. Morales.  1995.  The advertisement call of the leptodactylid frog Leptodactylus griseigularis.  Amphibia-Reptilia, 16:91-92.

Heyer, W. R., & J. A. Peters.  1971.  The frog genus Leptodactylus in Ecuador.  Proceedings of the Biological Society of Washington, 84:163-170.

Heyer, W. R., & W. F. Pyburn.  1983.  Leptodactylus riveroi, a new frog species from Amazonia, South America (Anura: Leptodactylidae).  Proceedings of the Biological Society of Washington, 96:560-566.

Heyer, W. R., & A. S. Rand. 1977. Foam nest construction in the leptodactylid frogs Leptodactylus pentadactylus and Physalaemus pustulosus (Amphibia, Anura, Leptodactylidae). Journal of Herpetology, 11:225-228.

Heyer. W. R., A. S. Rand, C. A. G. Cruz, O. L. Peixoto, & C. E. Nelson. 1990.  Frogs of Boracéia.  Arquivos de Zoologia, 31:231-410. 

Heyer, W. R., & Y. R. Reid. 2003. Does advertisement call variation coincide with genetic variation in the genetically diverse frog taxon currently known as Leptodactylus fuscus (Amphibia: Leptodactylidae)? Anais da Academia Brasileira de Ciências, 75:39-54.

Heyer, W. R., & N. J. Scott, Jr.  2006.  The advertisement call of Leptodactylus laticeps (Amphibia, Anura, Leptodactylidae):  Predatory aural luring?  Herpetological Natural History, 9(2):189-194.

Heyer, W. R., & P. A. Silverstone.  1969.  The larva of Leptodactylus hylaedactylus (Amphibia, Leptodactylidae).  Fieldiana: Zoology, 51:141-145.

Heyer, W. R., & A. S. Thompson.  2000.  Leptodactylus rugosus.  Catalogue of American Amphibians and Reptiles, (708):1-5.

Klappenbach, M. A., & J. A. Langone.  1992.  Lista sistemática y sinonímica de los anfibios del Uruguay.  Con comentarios y notas sobre su distribución.  Anales del Museo Nacional de Historia Natural de Montevideo, Segunda Série, 8:163-222.

Langone, J. A.  1995 ("1994").  Ranas y sapos del Uruguay (reconocimiento y aspectos biológicos).  Museo Damaso Antonio Larrañaga, Serie de Divulgación, (5):1-123.

Langone, J.A. and R. O. de Sá.  2005.  Redescripción de la morfología larval externa de dos especies del grupo de Leptodactylus fuscus (Anura, Leptodactylidae).  Phyllomedusa, 4:49-59.

Larson, P. M., & R. O. de Sá. 1998.  Chondrocranial morphology of Leptodactylus larvae (Leptodactylidae: Leptodactylinae):  Its utility in phylogenetic reconstruction.  Journal of Morphology, 238:287-305.

Larson, P. M., R. O. de Sá, & D. Arrieta.  2003.  Chondrocranial, hyobranchial and internal oral morphology in larvae of the basal bufonid genus Melanophryniscus (Amphibia: Anura).  Acta Zoologica (Stockholm), 84:145-154.

Lavilla, E. O., J. A. Langone, U. Caramaschi, W. R. Heyer, and R. O. de Sá.  2010.  The identification of Rana ocellata Linnaeus, 1758.  Nomenclatural impact on the species currently known as Leptodactylus ocellatus (Leptodactylidae) and Osteopilus brunneus (Gosse, 1851) (Hylidae).  Zootaxa, (2346):1-16. Send e-mail request to rdesa@richmond.edu or heyerr@si.edu for a pdf file of this paper.

Maxson, L. R., & W. R. Heyer.  1988.  Molecular systematics of the frog genus Leptodactylus (Amphibia: Leptodactylidae).  Fieldiana:  Zoology, New Series, (41):1-13.

Muedeking, M. H., & W. R. Heyer. 1976. Descriptions of eggs and reproductive patterns of Leptodactylus pentadactylus (Amphibia: Leptodactylidae). Herpetologica, 32:137-139.

Núñez, D., R. Maneyro, J. A. Langone, & R. O. de Sá.  2004.  Distribución geográfica de la fauna de anfibios del Uruguay.  Smithsonian Herpetological Information Service (134):1-34.

Padial, J. M., R. O. de Sá, & I. J. De la Riva.  2006.  The distress calls of Leptodactylus chaquensis Cei, 1950 and Leptodactylus elenae Heyer, 1978 (Anura: Leptodactylidae).  Salamandra, 42:57-60.

Ponssa, M. L., & W. R. Heyer.  2007.  Osteological characterization of four putative species of the genus Adenomera (Anura: Leptodactylidae), with comments on intra- and interspecific variation.  Zootaxa, (1403):37-54.

Pyburn, W. F., & W. R. Heyer.  1975.  Identity and call of the frog, Leptodactylus stenodema.  Copeia, 1975(3):585-587.

Radice, G., M. K. Boggiano, M. DeSantis, P. M. Larson, J. Oppong, M. Smetanick, T. Stevens, J. Tripp, R. Weber, M. Kerckhove, & R. O. de Sá.  1999.  Three-dimensional reconstructions of tadpole chondrocrania from histological sections.  Virginia Journal of Science, 50:227-235.

Straughan, I. R., & W. R. Heyer.  1976.  A functional analysis of the mating calls of the neotropical frog genera of the Leptodactylus complex (Amphibia, Leptodactylidae).  Papéis Avulsos de Zoologia, 19(23):221-245.

Vanzolini, P. E., & W. R. Heyer.  1985.  The American herpetofauna and the interchange.  Pages 475-487 in F. G. Stehli & D. S. Webb (eds.), The Great American Biotic Interchange.  Plenum Press.

Vaz-Ferreira, R., R. O. de Sá, F. Achaval, & A. Gehrau.  1984.  Leptodactylus podicipinus (Cope, 1862) y Leptodactylus chaquensis Cei, 1950 (Anura, Leptodactylidae), en el Uruguay.  Boletín de la Sociedad Zoológica del Uruguay, Segunda Época, 2:72-77.

Wassersug, R. J., & W. R. Heyer.  1988.  A survey of internal oral features of leptodactylid larvae (Amphibia: Anura).  Smithsonian Contributions to Zoology, (457):1-99.  [This article is broken up into four pdf files.  pdf file 1.  pdf file 2.  pdf file 3.  pdf file 4.]

Wynn, A., & W. R. Heyer.  2001.  Do geographically widespread species of tropical amphibians exist?  An estimate of genetic relationships within the Neotropical frog Leptodactylus fuscus (Schneider 1799) (Anura Leptodactylidae).  Tropical Zoology, 14:255-285.

Yanek, K., W. R. Heyer, & R. O. de Sá.  2006. Genetic resolution of the enigmatic Lesser Antillean distribution of the frog Leptodactylus validus (Anura, Leptodactylidae).  South American Journal of Herpetology, 1(3):192-201.

 

 
 

 

 
 
 
 
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